User Contributed Dictionary
Noun
cetaceans- Plural of cetacean
Extensive Definition
The order Cetacea (, L. cetus, whale) includes
whales, dolphins and porpoises. Cetus is Latin and is used in
biological names to mean "whale"; its original meaning, "large sea
animal," was more general. It comes from Ancient
Greek κῆτος (kētos), meaning "whale" or "any huge fish or
sea
monster". Cetology is the
branch of marine science associated with the study of
cetaceans.
Cetaceans are the mammals most fully adapted to
aquatic life. Their body is fusiform (spindle-shaped). The
forelimbs are modified into flippers. The tiny hindlimbs are
vestigial; they do not
attach to the backbone and are hidden within the body. The tail has
horizontal flukes. Cetaceans are nearly hairless, and are insulated
by a thick layer of blubber. As a group, they are
noted for their high
intelligence.
The order Cetacea contains ninety species, all
marine except for five species of freshwater dolphins. The order is
divided into two suborders, Mysticeti (baleen
whales) and Odontoceti
(toothed whales, which includes dolphins and porpoises).
Respiration, vision, hearing and echolocation
As mammals, cetaceans need to breathe air.
Because of this, they need to come to the water's surface to exhale
carbon
dioxide and inhale a fresh supply of oxygen. During diving, a muscular
action closes the blowholes
(nostrils), which remain closed until the cetacean next breaks the
surface; when it surfaces, the muscles open the blowholes and warm
air is exhaled.
Cetaceans' blowholes have
evolved to a position on top of the head, allowing more time to
expel stale air and inhale fresh air. When the stale air, warmed
from the lungs, is exhaled, it condenses as it meets the cold air
outside. As with a terrestrial mammal breathing out on a cold day,
a small cloud of 'steam' appears. This is called the 'blow' or
'spout' and is different in terms of shape, angle and height, for
each cetacean species. Cetaceans can be identified at a distance,
using this characteristic, by experienced whalers or
whale-watchers.
Cetaceans can go underwater for much longer
periods of time than other mammals. Their duration under water
varies greatly between speicies due to large physiological
differences between many members of this Order. There are two
studied advantages of cetacean physiology that let this Order (and
other marine mammals) forage underwater for extended periods of
time without breathing at the water surface.
Myoglobin concentrations in skeletal muscle of
mammals have much variation. A New Zealand white rabbit has
0.08+/-0.06 g (in a 100 g Wet muscle) of myoglobin, whereas a Northern
bottlenose whale has 6.34 g (in a 100 g Wet muscle) of myoglobin.
Myoglobin, by nature, has a higher affinity to oxygen than haemoglobin. That is,
myoglobin retains oxygen molecules better than hemoglobin.
Therefore, it is useful to have higher concentrations of myoglobin
when needed and there is no oxygen available for re-uptake. The
higher the myoglobin concentration in cetacean skeletal muscle, the
longer they can stay underwater and forage.
Increased body size is another way of elongating
dive duration of large cetaceans. This is true because of two
considered aspects. An increase in body size means that there is
increase in muscle mass, therefore, increase in muscle oxygen
stores. Another aspect is the universal correlation of mass and
metabolic rate (Kleiber's
law). In layman’s terms Kleiber’s law states that the metabolic
rate of a large animal is slower than a small animal per unit mass.
From this we can conclude that larger animals will use up less
oxygen than smaller animals (per mass unit).
The cetacean's eyes are set well back and to
either side of its huge head. This means that cetaceans with
pointed 'beaks' (such as dolphins) have good binocular vision
forward and downward but others, with blunt heads (such as the
Sperm Whale), can see either side but not directly ahead or
directly behind. Tear glands secrete greasy tears, which protect
the eyes from the salt in the water. Cetaceans also have an almost
spherical lens in their eyes, which is most efficient at focusing
what little light there is in the deep waters. Cetaceans make up
for their generally quite poor vision (with the exception of the
dolphin) with excellent hearing.
As with the eyes, the cetacean's ears are also
small. Life in the sea accounts for the cetacean's loss of its
external ears, whose function is to collect airborne sound waves
and focus them in order for them to become strong enough to hear
well. However, water is a better conductor of sound than air, so
the external ear was no longer needed: It is no more than a tiny
hole in the skin, just behind the eye. The inner ear, however, has
become so well developed that the cetacean can not only hear sounds
dozens of miles away, but it can also discern from which direction
the sound comes.
Some cetaceans are capable of echolocation.
Many toothed whales emit clicks similar to those in echolocation,
but it has not been demonstrated that they echolocate. Mysticeti
have little need of echolocation, as they prey upon small fish that
would be impractical to locate with echolocation. Some members of
Odontoceti, such as dolphins and porpoises, do perform
echolocation. These cetaceans use sound in the same way as bats -
they emit a sound (called a click), which then bounces off an
object and returns to them. From this, cetaceans can discern the
size, shape, surface characteristics and movement of the object, as
well as how far away it is. With this ability cetaceans can search
for, chase and catch fast-swimming prey in total darkness.
Echolocation is so advanced in most Odontoceti that they can
distinguish between prey and non-prey (such as humans or boats);
captive cetaceans can be trained to distinguish between, for
example, balls of different sizes or shapes.
Cetaceans also use sound to communicate, whether
it be groans, moans, whistles, clicks or the complex 'singing' of
the Humpback
Whale.
Feeding
When it comes to food and feeding, cetaceans can
be separated into two distinct groups. The toothed whales,
Odontoceti like sperm whales, beluga whales, dolphins and
porpoises, usually have lots of teeth that they use for catching
fish, squid or other marine life. They do not chew their food, but
swallow it whole. In the rare cases that they catch large prey, as
when Orca (Orcinus orca) catch a seal, they tear chunks off it that
in turn are swallowed whole.
The baleen whales or Mysticeti do not have teeth.
Instead they have plates made of keratin (the same substance as
human fingernails) which hang down from the upper jaw. These plates
act like a giant filter, straining small animals (such as krill and fish) from the seawater.
Cetaceans included in this group include the Blue Whale, the
Humpback Whale, the Bowhead Whale and the Minke Whale.
Not all Mysticeti feed on plankton: the larger
whales tend to eat small shoaling fish, such as herrings and
sardine, called micronecton. One species of Mysticeti, the Gray Whale
(Eschrichtius robustus), is a benthic feeder,
primarily eating sea floor crustaceans.
Mammalian nature
Cetaceans are mammals, that is, members of the
class
Mammalia.
The closest living relative of cetaceans is the hippopotamus.
As mammals, cetaceans have characteristics that
are common to all mammals: They are warm-blooded,
breathe in air through their lungs, bear their young alive and
suckle them on their own milk, and have hair, although very little of
it.
Another way of discerning a cetacean from a fish
is by the shape of the tail. The tail of a fish is vertical and
moves from side to side when the fish swims. The tail of a cetacean
– called a fluke – is horizontal and moves up and down, as
cetaceans' spines bend in the same manner as a human spine.
Taxonomic listing
The classification here closely follows Dale W.
Rice, Marine Mammals of the World: Systematics and Distribution
(1998), which has become the standard taxonomy reference in the
field. There is very close agreement between this classification
and that of Mammal Species of the World: 3rd Edition (Wilson and
Reeder eds., 2005). Any differences are noted using the
abbreviations "Rice" and "MSW3" respectively. Further differences
due to recent discoveries are also noted.
Discussion of synonyms and subspecies are
relegated to the relevant genus and species articles.
- ORDER CETACEA
- Suborder Mysticeti: Baleen
whales
- Family Balaenidae:
Right
whales and Bowhead
Whale
- Genus Balaena
- Bowhead Whale, Balaena mysticetus
- Genus Eubalaena
- North Atlantic Right Whale, Eubalaena glacialis
- North Pacific Right Whale, Eubalaena japonica
- Southern Right Whale, Eubalaena australis
- Genus Balaena
- Family Balaenopteridae:
Rorquals
- Subfamily Balaenopterinae
- Genus Balaenoptera
- Common Minke Whale, Balaenoptera acutorostrata
- Antarctic Minke Whale, Balaenoptera bonaerensis
- Sei Whale, Balaenoptera borealis
- Bryde's Whale, Balaenoptera brydei
- Eden's Whale Balaenoptera edeni - Rice lists this as a separate species, MSW3 does not
- Balaenoptera omurai - MSW3 lists this is a synonym of Bryde's Whale but suggests this may be temporary.
- Blue Whale, Balaenoptera musculus
- Fin Whale, Balaenoptera physalus
- Genus Balaenoptera
- Subfamily Megapterinae
- Genus Megaptera
- Humpback Whale, Megaptera novaeangliae
- Genus Megaptera
- Subfamily Balaenopterinae
- † Genus Eobalaenoptera
- † Harrison's Whale, Eobalaenoptera harrisoni
- Family Eschrichtiidae
- Genus Eschrichtius
- Gray Whale, Eschrichtius robustus
- Genus Eschrichtius
- Family Neobalaenidae:
Pygmy Right Whale
- Genus Caperea
- Pygmy Right Whale, Caperea marginata
- Genus Caperea
- Family Balaenidae:
Right
whales and Bowhead
Whale
- Suborder Odontoceti:
toothed whales
- Family Delphinidae:
Dolphin
- Genus Cephalorhynchus
- Commerson's Dolphin, Cephalorhyncus commersonii
- Chilean Dolphin, Cephalorhyncus eutropia
- Heaviside's Dolphin, Cephalorhyncus heavisidii
- Hector's Dolphin, Cephalorhyncus hectori
- Genus Delphinus
- Long-beaked Common Dolphin, Delphinus capensis
- Short-beaked Common Dolphin, Delphinus delphis
- Arabian Common Dolphin, Delphinus tropicalis. Rice recognises this as a separate species. MSW3 does not.
- Genus Feresa
- Pygmy Killer Whale, Feresa attenuata
- Genus Globicephala
- Short-finned Pilot Whale, Globicephala macrorhyncus
- Long-finned Pilot Whale, Globicephala melas
- Genus Grampus
- Risso's Dolphin, Grampus griseus
- Genus Lagenodelphis
- Fraser's Dolphin, Lagenodelphis hosei
- Genus Lagenorhynchus
- Atlantic White-sided Dolphin, Lagenorhynchus acutus
- White-beaked Dolphin, Lagenorhynchus albirostris
- Peale's Dolphin, Lagenorhynchus australis
- Hourglass Dolphin, Lagenorhynchus cruciger
- Pacific White-sided Dolphin, Lagenorhynchus obliquidens
- Dusky Dolphin, Lagenorhynchus obscurus
- Genus Lissodelphis
- Northern Right Whale Dolphin, Lissodelphis borealis
- Southern Right Whale Dolphin, Lissodelphis peronii
- Genus Orcaella
- Irrawaddy Dolphin, Orcaella brevirostris
- Australian Snubfin Dolphin, Orcaella heinsohni. 2005 discovery, thus not recognized by Rice or MSW3 and subject to revision.
- Genus Orcinus
- Killer Whale, Orcinus orca
- Genus Peponocephala
- Melon-headed Whale, Peponocephala electra
- Genus Pseudorca
- False Killer Whale, Pseudorca crassidens
- Genus Sotalia
- Tucuxi, Sotalia fluviatilis, but see the species article for a discussion
- Genus Sousa
- Pacific Humpback Dolphin, Sousa chinensis
- Indian Humpback Dolphin, Sousa plumbea
- Atlantic Humpback Dolphin, Sousa teuszii
- Genus Stenella
- Pantropical Spotted Dolphin, Stenella attenuata
- Clymene Dolphin, Stenella clymene
- Striped Dolphin, Stenella coeruleoalba
- Atlantic Spotted Dolphin, Stenella frontalis
- Spinner Dolphin, Stenella longirostris
- Genus Steno
- Rough-toothed Dolphin, Steno bredanensis
- Genus Tursiops - Rice
and MSW3 tentatively agree on this classification but see species
article for more detail.
- Indian Ocean Bottlenose Dolphin, Tursiops aduncus
- Common Bottlenose Dolphin, Tursiops truncatus
- Genus Cephalorhynchus
- Family Monodontidae
- Genus Delphinapterus
- Beluga, Delphinapterus leucas
- Genus Monodon
- Narwhal, Monodon monoceros
- Genus Delphinapterus
- Family Phocoenidae:
Porpoises
- Genus Neophocaena
- Finless Porpoise, Neophocaena phocaenoides
- Genus Phocoena
- Spectacled Porpoise, Phocoena dioptrica
- Harbour Porpoise, Phocoena phocaena
- Vaquita, Phocoena sinus
- Burmeister's Porpoise, Phocoena spinipinnis
- Genus Phocoenoides
- Dall's Porpoise, Phocoenoides dalli
- Genus Neophocaena
- Family Physeteridae:
Sperm
Whale family
- Genus Physeter
- Sperm Whale, Physeter catodon (syn. P. macrocephalus)
- Genus Physeter
- Family Kogiidae - MSW3
treats Kogia as a member of Physeteridae
- Genus Kogia
- Pygmy Sperm Whale, Kogia breviceps
- Dwarf Sperm Whale, Kogia sima
- Genus Kogia
- Superfamily Platanistoidea:
River dolphins
- Family Iniidae
- Genus Inia
- Amazon River Dolphin, Inia geoffrensis
- Genus Inia
- Family Lipotidae - MSW3 treats Lipotes as a member of Iniidae
- Family Pontoporiidae
- MSW3 treats Pontoporia as a member of Iniidae
- Genus Pontoporia
- Franciscana, Pontoporia blainvillei
- Genus Pontoporia
- Family Platanistidae
- Genus Platanista
- Ganges and Indus River Dolphin, Platanista gangetica. MSW3 treats Platanista minor as a separate species, with common names Ganges River Dolphin and Indus River Dolphin, respectively.
- Genus Platanista
- Family Iniidae
- Family Ziphidae, Beaked
whales
- Genus Berardius
- Arnoux's Beaked Whale, Berardius arnuxii
- Baird's Beaked Whale (North Pacific Bottlenose Whale), Berardius bairdii
- Subfamily Hyperoodontidae
- Genus Hyperoodon
- Northern Bottlenose Whale, Hyperoodon ampullatus
- Southern Bottlenose Whale, Hyperoodon planifrons
- Genus Indopacetus
- Indo-Pacific Beaked Whale (Longman's Beaked Whale), Indopacetus pacificus
- Genus Mesoplodon,
Mesoplodont
Whale
- Sowerby's Beaked Whale, Mesoplodon bidens
- Andrews' Beaked Whale, Mesoplodon bowdoini
- Hubbs' Beaked Whale, Mesoplodon carlhubbsi
- Blainville's Beaked Whale, Mesoplodon densirostris
- Gervais' Beaked Whale, Mesoplodon europaeus
- Ginkgo-toothed Beaked Whale, Mesoplodon ginkgodens
- Gray's Beaked Whale, Mesoplodon grayi
- Hector's Beaked Whale, Mesoplodon hectori
- Layard's Beaked Whale, Mesoplodon layardii
- True's Beaked Whale, Mesoplodon mirus
- Perrin's Beaked Whale, Mesoplodon perrini. This species was recognised in 2002 and as such is listed by MSW3 but not Rice.
- Pygmy Beaked Whale, Mesoplodon peruvianus
- Stejneger's Beaked Whale, Mesoplodon stejnegeri
- Spade Toothed Whale, Mesoplodon traversii
- Genus Hyperoodon
- Genus Tasmacetus
- Tasman Beaked Whale (Shepherd's Beaked Whale), Tasmacetus shepherdi
- Genus Ziphius
- Cuvier's Beaked Whale, Ziphius cavirostris
- Genus Berardius
- Family Delphinidae:
Dolphin
- Suborder Mysticeti: Baleen
whales
See also
References
External links
- ARKive - images and movies of Whales, Dolphins and Porpoises
- Tree of Life illustration showing how Cetacea relates to other lifeforms
- images and movies of Whales and Dolphins. Text is in French
- http://www.acsonline.org/ American Cetacean Society
- British Cetacean Site especially interesting is taxonomy
- Cetacea.org homepage
- Walker's Mammals of the World Online - Cetaceans
- Tursiops.org: Current Cetacean-related news
- Dolphins and Whales on Science Daily
- All About Dolphins: Information and News Concerning Dolphins and other Cetacea
- http://www.sonsdemar.eu/ Sounds of the cetaceans, bioindicators, acoustic trauma, acoustic signals
- Cetacea Evolution - by Douglas J. Futuyma
cetaceans in Min Nan: Cetacea
cetaceans in Arabic: حيتانيات
cetaceans in Bulgarian: Китоподобни
cetaceans in Catalan: Cetaci
cetaceans in Czech: Kytovci
cetaceans in Danish: Hvaler
cetaceans in German: Wale
cetaceans in Estonian: Vaalalised
cetaceans in Modern Greek (1453-): Κητώδη
cetaceans in Spanish: Cetacea
cetaceans in Esperanto: Cetacoj
cetaceans in Persian: آببازان
cetaceans in French: Cetacea
cetaceans in Galician: Cetáceo
cetaceans in Korean: 고래
cetaceans in Hindi: व्हेल
cetaceans in Croatian: Kitovi
cetaceans in Indonesian: Cetacea
cetaceans in Icelandic: Hvalir
cetaceans in Italian: Cetacea
cetaceans in Hebrew: לווייתנאים
cetaceans in Latin: Cetacea
cetaceans in Lithuanian: Banginiai
cetaceans in Limburgan: Walvèsechtege
cetaceans in Hungarian: Cetek
cetaceans in Malay (macrolanguage): Ikan
paus
cetaceans in Dutch: Walvissen
cetaceans in Japanese: クジラ目
cetaceans in Norwegian: Hvaler
cetaceans in Norwegian Nynorsk: Kvalar
cetaceans in Polish: Walenie
cetaceans in Portuguese: Cetáceos
cetaceans in Romanian: Cetaceu
cetaceans in Russian: Китообразные
cetaceans in Sicilian: Citacea
cetaceans in Simple English: Cetacea
cetaceans in Slovak: Veľryby (rad)
cetaceans in Slovenian: Kiti
cetaceans in Serbian: Китови
cetaceans in Finnish: Valaat
cetaceans in Swedish: Valar
cetaceans in Thai: อันดับวาฬและโลมา
cetaceans in Turkish: Balinalar
cetaceans in Contenese: 鯨魚
cetaceans in Chinese: 鯨目